Vibrissa well above lower margin of eye; head in profile rounded ventrally, distinctly higher than long. Vertical diameter of eye usually at least 1.5x the horizontal diameter. Frons in male narrower than in female. Wing with subcostal notch usually evident, sometimes strongly so, with basal sector of C ending in an acute lappet; R4+5 and M1 sometimes converging at tips; distal margin of anal cell meeting anal vein in sharp angle. Abdominal tergites of male often with silvery microtomentum.