Vibrissa below or at the level of lower margin of eye; head in profile subquadrate; vertical diameter of eye less than 1.5x the horizontal diameter; postocellar setae only slightly converging or parallel. Frons in male as wide as in female. Basisternum large and V-shaped, sometimes with a precoxal bridge. Wing with subcostal notch seldom evident, never well-developed; R4+5 and M1 not or only slightly converging at tips; distal margin of anal cell (CuA2) usually rounded (if not, then epistoma extended dorsally, subtriangular). First tarsomere of hind leg without a posteroventral comb of setae. Abdominal tergites of male without silvery microtomentum.